at least the users role.
people in this role are allowed to view the project.
depending on your permission scheme you could define very comfortable what project role members are able to do.
if you want to create special permissions then maybe its a good plan to create a new permission scheme for your project where you only want to consider project roles instead of user-groups like "jira-users, jira-developers, jira-administrators"
because every active user who is able to login to jira must be in "jira-users" the default permissions allow project view access to anybody in this group. to avoid this create the new scheme and assign it to your project. then remove groups from the new scheme in the roles tab and add only the persons you want.
try only editing the roles of your project. clear the groups in developers and users then add your users you want to access
for more details check https://confluence.atlassian.com/display/JIRA/Managing+Project+Permissions
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