I have tested in JIRA Core 7.2, that users not belonging to jira-users or admins and without an application can still receive email notifications if checkd as active.
I'd like to confirm
As long as the user has a valid email address, and can see the issue if they were to visit it in the UI, then yes, they can get the emails.
Any user who can log in counts towards your licence number (if you have Service Desk, then "customers" are unlimited, but that's a different type of login)
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