I have used Source Tree with the same repository and the same paths for months (in a Windows environment) then suddenly last Monday, this error has started to appear when the program is opening the repository.
I have already tried with the command line
git config --system core.longpaths true
to avoids the length of the paths in a Windows environment, but it does not wok.
But most important: why it has worked perfectly since the beginning and only now there is this problem?
Try git operations from the command line (fetch, checkout, branch, push, etc). If they have the same error, then you should take your question to a site like StackOverflow that has a larger number of git "power users", as compared to the UI-users that are subscribed to the "sourcetree" tag on this site.
If the command line works, then it sounds to me like you've got a candidate for a bug report at jira.atlassian.com.
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