For example, instead of https://<domain>.atlassian.net/servicedesk/customer/portal/2
can we make it something like
https://<domain>.atlassian.net/servicedesk/customer/portal/<projectname>
or even https://<domain>.atlassian.net/servicedesk/customer/<projectname>
Unfortunately it will not be possible for you to perform URL rewriting when you are hosted in the Cloud, in On-Demand. This is typically done by a system administrator or network administrator. They would start by modifying configurations of the Tomcat or Apache front end, which you do not have access to in this scenario.
Atlassian OnDemand Restricted Functions are listed here: https://confluence.atlassian.com/display/AOD/Restricted+Functions+in+Atlassian+OnDemand
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