There are several ways this can get into the logs, and you've only given us the error without any tracing information.
But, I've mostly seen this error from old versions of Agile - what version of Agile and JIRA are you using?
In JIRA i can see mails getting piled up in mail queue.
1)Mail server configuration is fine,2)User details are valid but still they get queued .
Error thrown in logs was something like this
vzhvzs 10.xx.xx.xxx /browse/TestAgent-658 [atlassian.plugin.webresource.DefaultResourceDependencyResolver] Cannot find web resource module for: com.atlassian.auiplugin:aui-experimental-tooltips.
No idea how to resolve this.
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