I've just noticed it there. It's not enabled currently and rather than just enable it, I'd like to know what it does first. All it says is "This plugin provides cross-product Quick Nav in a harmonious way". I've found no documentation on it and no mention of it on the internet.
This is a new feature that is currently in development. The feature enhances the quick search between the products. As the feature is still in development, there is currently no additional documentation available and the feature is disabled for your OnDemand instance.
You can enable it, but as mentioned there is currently no supporting documentation for the feature and it is still in development. If stability is a major concern for your instance I would recommend leaving this disabled until a formal release occurs.
The applicaiton navigator is a configurable drop-down menu that allows quick-access switching between connected Atlassian applications, or any other application you configure. Have a look here for the FAQ:
Hope this helps and thanks for posting on Answers!
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