It use to be when you queried Confluence via SOAP APIs (for instance, with getServerInfo) you got an accurate representation of the version you were attached to. Not any more:
Download: Confluence version: 5.4.1, build: 4727 OnDemand: Confluence version: 5.4.0, build: 4913
Version numbering on OnDemand is misleading since multiple features are added/removed compared to the Download version. Besides logins and applinks (which are pre-wired in OnDemand), think about the experiments that Atlassian runs and sometimes confirms and sometimes throws away.
Also, versions are used internally to compare many things, for example for imports/exports and the "What's New" dialog. It is not obvious at all for a developer what "should" be the OD version.
The only things which are wise to consider are:
Best of luck for whatever you're attempting to perform with those numbers.
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